Historical movement and cultivation amplified that blending. After early domestication in Asia, citrus spread through Austronesian movement and later westward trade routes into the Mediterranean, where additional selection and crossing produced familiar market groups. That is why sweet orange, grapefruit, lemon, and clementine are better understood as overlapping recombination clusters than as isolated branches. Over time, human breeding tended toward sweeter, more mandarin-rich market varieties - as can be seen in the path from pomelo to mandarin, which is largely unidirectional.
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